Why do the Duga-1 dipoles look like an array of birdcages?
This is a cage dipole antenna (or an array of cage dipoles to be exact). It works just like a dipole, but has a wider bandwidth. The thicker the dipole, the wider the bandwidth.
RF does not care if the conductors are hollow cylinders, solid surfaces, cages or mesh as long as size of the "holes" is signifficantly smaller than the wavelength (that is also why you can look into the microwave oven through tiny holes and not get cooked in the process).
That second image shows a feed where you would expect it. It's a bit hard to see on all of the elements, and it's better to look at the high resolution original image. But they're definitely center fed.
Specifically, see https://commons.m.wikimedia.org/wiki/File:Chernobyl_Exclusion_Zone_Antenna_hnapel_22.jpg and https://commons.m.wikimedia.org/wiki/File:Chernobyl_Exclusion_Zone_Antenna_hnapel_14.jpg for close-ups of the antenna feed connections (hooray for Wikimedia commons).
The birdcage construction is mainly to save material. The RF currents in the antenna only flow along the long axis of the antenna, so you just need enough parallel conductors in that direction. And due to the skin effect, the RF current only flows on the outer surface, so the center can be completely hollow. The circular metal pieces do not carry RF current and simply serve to strengthen the structure. It's quite common to see antennas with ground planes that are just a bunch of parallel wires. Speaking of planes, it looks like the antennas are located in front of a reflector, which appears to be made of a large number of thin horizontal wires. Provided the wires are sufficiently close together, this works exactly the same as a continuous sheet, but requires far less material and hence is less expensive, requires less support, and allowes wind and rain to pass through the antenna.
Now, as for the double cone shape, I'm not sure why that's necessary offhand. It's possible they're going for a biconical wideband element, but it seems to me like the array and reflector spacing would limit the overall bandwidth.